I'll way in. I've owned a 5th gen 4runner for the past 8 years. Had a brief stent with a 2018 TRD off-road road premium then in January 2029 bought my current TRD Pro brand new off the lot. Six plus years later and almost 120k miles, suffice it to say it is an EXCELLWNT VEHICLE. It's issues are well known. Slow. Not great fuel economy. Outdated tech. The tech is VERY easy to remedy. As to the other two, slow is no issue as it is not a race car. It's a go anywhere and get you back car. And with the fuel tank size, I never felt like the low gase milage was a problem at all.
My wife and I were VERY BLESSED to come into our Land Cruiser LC 250 w/Premium Package. It is not stock and has several professionaly installed aftermarket mods so when I discuss it keep that in mind. Having only had it since January and in light of the mods it's hard to make a fair comparison. But I will do so anyway. The LC at base is heavier than our 4runner, and with the mods rides much higher and is even heavier still. Despite that, it drives better, handles better, and while no race car, is still noticeably faster than the 4runner and noticeably quicker off the line.
The interior is most definitely nicer/more plush as well. And the tech is light years better than the 4runner came with stock. Interior space (not including the trunk space) is also better in the LC than the 4runner. Trunk space wise it seems to be a wash to me. Some may include this with tech, but in terms of safety features, the LC is loaded and makes my wife and I feel safer while driving. This is a big deal considering we have our two small children riding in the back with us.
Road and wind noise/interior sound deadening wise even with the tent and rack onto and bigger heavier tires, LC 250 is better/quiter. Sound system wise, both are JBL systems, and Landcruiser is just better. Hands down.
As for fuel economy, at base I am very confident that the LC 250 is better. However with the 35s, 3 inch lift, all the added weight, plus the RTT on top that acts like an air damn, the fuel economy on the LC now is, shall we say, less than stellar. Combined with the small fuel tank, and it is a serious Achilles heel. Womp womp.
The one thing that I will give the 4runner, and is just not yet known with this new Lancruiser 250, is that I never doubt at all if my 4runner is going to work, if it's going to break down, if I'm going to have problems with the truck. It is a TANK of confidence in that way. The LC, I hope it will be that way. But it's a question mark for me.
So far though my wife and I are very impressed.
Some will say hey, your comparing it to a 4runner! You should compare to the old LC 200!! Well, I haven't driven that one. My 4runner is the top trim of its generation. And just as, if not more, reliable than the 200 is. And definitely more capable off road than the 200 is though the 200 is no chump off road. This LC 250 we have (yes heavily molded for off road) is unquestionably better off road than our 4runner TRD Pro is.
Let's look at some hard facts between 250 and 200. 200 is faster, slightly and has more HP. 250 is lighter, has better fuel economy, and has more torque. 200 can tow more (by a about a ton!). 250 is verifiable better off road. At base, 250 has significantly better tech than the 200 and has better safety features. 200 has a 3rd row, 250 doesn't. Oh and starting price. Base 200 I think in it's last year was more than a 250 FE. So price wise the 250 is LESS expensive (a plus in my book) so if you add those up, it's a pretty lopsided win for the 250. The one big x factor is longevity and reliability. Which we just don't know yet on the 250. Only time will tell
The last thing I will say, there is not a single 200 LC out there, modded or otherwise, that i would trade for our 250. Period.